Torus (Doughnuts)
GFP is targeted to the lumen of numerous small organelles. The labeled organelles vary in shape, frequently adopting toroid and tubular forms that can interconvert. The "hole" in the torus may either be a true hole, an exclusion of GFP from the center of non-perforated organelle, or a dimple such as that in a red blood cell. Sequence analysis of the tags, and in situ analysis suggest that these organelles are peroxisomes (sequence and in situ analysis).
Variable torus organelle morphology and motility
Interconversion of forms
Tubule extension and retraction
Clustering and tubule anchoring

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Organelle dynamics 
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